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She sometimes appeared in sketches on Late Night with Conan O'Brien.She had a recurring role on Karen Sisco as Detective Marley Novak in 20.In 2005, she was cast in her breakout role in the ABC series Grey's Anatomy, as Dr.Addison Montgomery, the estranged spouse of Derek Shepherd ("Dr. It was announced in February 2007 that Walsh's Grey's Anatomy character would be in a spin-off of Grey's Anatomy that began airing in September 2007.In 2014, Walsh had a supporting role in the first season of FX's television series Fargo.She had a guest starring role in Hulu's original series The Hotwives of Orlando and later starred in the NBC series Bad Judge, which aired for one season.After playing Cathy Buxton in the Homicide: Life on the Street episode "Stakeout" in 1996 and Navy Lieutenant Kirstin Blair in the Law & Order episode "Navy Blues", her first major television appearance was in 1997, when she appeared on The Drew Carey Show as Drew Carey's love interest, Nicki Fifer.She wore a fatsuit in some episodes of the show, as the Nicki character was a formerly obese woman who lost weight and slowly started to gain it back.

Her performance was dubbed to be the "best work" of her career and was met with critical acclaim.

In New York, Walsh was a member of the improvisational comedy troupe Burn Manhattan and the New York City Players theater ensemble.

Launched via various social media channels, including Twitter, You Tube and select blogs, the Boyfriend webimercials have been viewed more than 200,000 times to date and were released by Walsh prior to her appearance on HSN to drive excitement and anticipation.

Walsh ran as a candidate with the Unite for Strength party who won six of the 11 Hollywood Division seats which were up for contention in the recent Screen Actor's Guild National Board elections.

In 2005, Walsh joined the Planned Parenthood Federation of America (PPFA)'s Board of Advocates, and in 2008 received the PPFA Maggie Award for an Artist who Stands Up for Reproductive Health and Freedom.

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